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Gregor Townsend may count the cost of injuries from Scotland's final Rugby World Cup warm-up victory over Georgia.
One of the best wings in South African rugby history, World Cup winner Chester Williams died on Friday.
Japan were unable to repeat their shock 2015 Rugby World Cup success over South Africa, largely due to Makazole Mapimpi's hat-trick.
The upcoming Rugby World Cup will mark David Pocock's last matches for the Wallabies.
Ireland have been boosted by the returns of Johnny Sexton, Conor Murray, Keith Earls and Robbie Henshaw for their meeting with Wales.
Wales have rung the changes against Ireland as they finish preparations for the Rugby World Cup.
Though he is not in the Rugby World Cup squad, Joe Marchant will start at centre against Italy in England's final warm-up game.
Set to make his 90th appearance for the Wallabies, veteran hooker Tatafu Polota-Nau is retiring from Tests at year's end.
Ireland's talisman Johnny Sexton says he was "blessed' to overcome a thumb injury in time to make Joe Schmidt's Rugby World Cup squad.
Hooker Stuart McInally has been selected to captain Scotland in Japan, though Huw Jones was the shock absentee from the squad list.
